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We offer a Physician Scientist training program for the fulfillment of requirements for MD and PhD degrees. The comprehensive program includes two years in pre-clinical medical training, followed by graduate training in one of our basic research departments and two years of medical rotations for clinical training.
The small size of the program ensures a high quality training experience providing individualized attention to its students. As a MD-PhD student you will experience great flexibility in designing a research program and have exceptional access to a wide variety of research experiences in top-flight research laboratories. You will receive hands-on experience in the patient simulation center and work with standardized patients beginning in the first year. A wide variety of clinical experiences are available through our hospital partnerships in Louisville and throughout Kentucky.
Applicants should have fulfilled prerequisites for admission to the School of Medicine - including the Medical College Admission Test and have a proven commitment to biomedical research.
Students in the program will receive full tuition remission, a stipend during all phases of the curriculum and health insurance during the graduate research phase.
Application and Interview Process
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Application Process
Applicants to the combined MD-PhD Program must submit applications to the Medical School through the American College Application Service (AMCAS) and the student is required to take the MCAT. If you are interested in being considered for the MD-PhD Program, you should check the appropriate box on the University of Louisville Medical School application when you apply. Checking this box will not influence admission to medical school and interested medical school students are encouraged to explore this option.
The MCAT application deadline is in August.
The Medical School AMCAS application deadline is Saturday, November 1, 2025.
The University of Louisville School Of Medicine deadline for the secondary application is Monday, December 1, 2025.
After initial screening of applications, selected candidates who have indicated an interest in the MD-PhD Program will be invited to interview with both the School of Medicine Admissions Committee and members of the MD-PhD Program Committee. Applicants will be notified by email. Emails will be sent to the primary email listed on your AMCAS application.
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The Combined MD-PhD Program Application Checklist
University of Louisville, Medical School Admissions:
Abell Administration Center
Louisville, KY 40202-3866
502-852-5193
Email: medadm@louisville.edu
Admissions Website
American Medical College Application Service:
Association of American Medical Colleges (AAMC)
Section for Student Services
2501 M Street, NW Lobby-26
Washington, DC 20037-1300
Email: medadm@louisville.edu
Website: www.aamc.org
Medical College Admissions Test Scores: To be considered for fall admission you should take the MCAT no later than fall prior to the year for which you wish to gain admission. MCAT scores more than two years old are not acceptable. Information regarding the MCAT can be obtained from:
MCAT Registration
The American College Testing Program
P.O. Box 414, Iowa City, IA 52243
319-337-1276
Email: mcat@aamc.org
Website: www.aamc.org/students/mcat
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Interview Process
Interviews will be conducted in-person and virtually for the 2026-2027 admissions cycle on multiple sets of dates. These interview dates are still being determined for the months of October and November.
Detailed instructions will be provided. The first day will be with the Medical School Admissions committee and the second day will be with members of the MD-PhD committee.
Research experience is an important aspect of the MD-PhD interviews and you should be prepared to discuss why you would like to be a physician-scientist, along with your interests and goals. Upon completion of the interviews, your application and interview evaluations will be reviewed by the MD-PhD Advisory Committee. Follow-up and feedback information will be made available to MD-PhD applicants in mid-December. Final candidate selection should be made by April of the matriculation year. If you are not offered a position in the MD-PhD Program, you must contact the School of Medicine to let them know if you want to be considered for the Regular MD program only.
Current students in the University of Louisville School of Medicine are eligible to join the MD-PhD Program after the completion of their second year of Medical School. Interested students should contact either of the co-directors of the MD-PhD Program: Maxwell Boakye or Kenneth Palmer, in the spring semester of their first year. This is an internal application process and selected applicants may be invited to interview with the MD-PhD Committee.
Additional Information
MD-PhD Program Co-Directors
The MD-PhD Program co-directors, Maxwell Boakye and Kenneth Palmer, bring complementary expertise in translational research and a shared commitment to scholarship, training and mentorship. Both have been recognized as the University of Louisville's “Student Champions” for their dedication to student success.
Maxwell Boakye, MD, MPH, MBA, FACS, FAANS
Dr. Boakye, a neurosurgeon-scientist with deep networks in the UofL Health - UofL Hospital clinical enterprise, complements this expertise by integrating clinical investigation skills into graduate training for students pursuing highly competitive specialties. Boakye’s laboratory research on porcine models of spinal cord injury complements his clinical research in neuromodulation to improve SCI outcomes. He published over 220 peer-reviewed manuscripts, with more than 8,000 citations and an H-index of 50 and holds certifications in healthcare improvement (IHI), patient safety (NAHQ) and Health Professions Education, equipping him with expertise in adult education and mentoring. He completed leadership programs at Harvard University, Cornell University and University of Louisville.
Kenneth E. Palmer, PhD
Dr. Palmer brings extensive experience as the director of the Center for Predictive Medicine for Biodefense and Emerging Infectious Diseases, including University of Louisville’s Regional Biocontainment Laboratory. He has been the Principal Investigator of complex multi-project NIH and DOD grants, including serving as Investigator-Sponsor of two first-in-human clinical trials, overseeing budget management and resource allocation. He has completed CIMER mentoring training. With service on over 20 PhD dissertation committees and successful mentoring of PhD students, a current MD-PhD student, postdoctoral trainees and junior faculty, Dr. Palmer, a first-generation college graduate, leads the graduate research training phase of the MD-PhD Program.
The School of Medicine has entered a phase of dynamic growth stimulated by the addition of space in state-of-the-art research buildings and the recruitment of a new generation of endowed research professors through the Kentucky Commonwealth's Bucks for Brains program. This growth has resulted in an increase in research funding over the past few years.
The school has targeted multiple areas in molecular medicine and biotechnology for scientific excellence, including therapeutic oncology and tumor cell biology, cardiovascular disease research, transplantation and immunology, neurobiology, aging, ophthalmology and visual sciences and contemporary issues in public health and bioethics. Our scientific investigators are working to develop new strategies to combat common diseases in our society, such as breast and lung cancer, sickle-cell anemia, AIDS, diabetes, digestive diseases, paralysis caused by spinal-cord injury and childhood academic problems caused by sleep disorders. New multidisciplinary centers have been established to study and identify mechanisms to decrease heart damage after a heart attack and to support cardiovascular function with internal artificial hearts and stimulate spinal cord regeneration with neural stem cells. University of Louisville also has a prominent team researching the genetics of aging using microarray technology and an internationally recognized group pioneering hand transplantation.
These are only a few of the exciting research projects that are underway at the University of Louisville. Please contact us and share our enthusiasm.
Basic Science Departments
- Anatomical Sciences and Neurobiology
- Biochemistry and Molecular Genetics
- Microbiology and Immunology
- Pharmacology and Toxicology
- Physiology
Research Opportunities
The School of Medicine at the University of Louisville offers many outstanding research opportunities in which MD-PhD students can obtain research training and excellent credentials. Examples of opportunities include, but are not limited to:
- Cardiovascular Research
- Cancer Research
- Cellular Therapeutics
- Diabetes and Obesity
- Digestive Health Research
- Spinal Cord Injury Research
- Vaccine Development for emerging, neglected or rare infectious diseases
- Vision Research and Ocular Disease
Research Mentors
MD-PhD students have the opportunity to complete a rotation prior to and after their first year of Medical School within laboratories they are considering for their dissertation. This allows the student to become familiar with the lab environment, the research projects and the mentor, before committing to the laboratory and dissertation mentor. If the mentor is associated with more than one graduate department, you will need to determine which department would be best for you to reach your goals.
The program combines pre-clinical coursework, graduate coursework and fundamental biomedical research resulting in a dissertation and a PhD degree and clinical training which will result in an MD degree. These two approaches emphasize the interface between biomedical science and clinical practice that is necessary in treating patients in tomorrow's world and investigating the basis of their disease. The program is outlined below:
SUMMER RESEARCH prior to Medical School Year One - up to 10 weeks
Medical School Year One (M1):
Clinical Anatomy, Development and Physical Examination (CADE): 19-week course in fall semester that integrates Gross Anatomy, Embryology and Physical Examination content by body region.
Molecular Basis of Life, Defense and Disease: 16-week course in spring semester that integrates foundational biochemistry, medical genetics, immunology, tissue and body fluid physiology, pathology and pharmacology to introduce immunologic function in disease and health; normal cellular and tissue histology in health, injury and healing; foundational microbiology; and mechanisms, diagnosis and treatment of neoplastic and hematologic diseases.
Introduction to Biostatistics and Evidence-Based Medicine: Two week course in spring semester that concentrates principles of biomedical research, foundational statistics, medical informatics and application of evidence to patient care.
Introduction to Clinical Medicine 1 (ICM 1): Two semester course that covers foundational patient interviewing skills, professionalism and ethics, humanism and compassion, medical system structure and health disparities, cultural humility, provision of health maintenance and use of health guidelines.
SUMMER RESEARCH in between MS Year One and MS Year Two - 10 weeks
Medical School Year Two (M2):
Systems in Health and Disease 1: 19-week course in fall semester that integrates the normal physiology of the cardiovascular, respiratory, renal, gastrointestinal, hepatobiliary, nutritional/metabolic and endocrine systems with their pathophysiologic function in disease, progression from normal histologic appearance to pathologic appearance, infecting microbes and applicable pharmacologic treatments.
Systems in Health and Disease 2: 14-week course in spring semester that integrates the normal physiology of the reproductive, dermatologic, musculoskeletal and behavioral systems as well as neurologic system diseases with their pathophysiologic function in disease, progression from normal histologic appearance to pathologic appearance, infecting microbes and applicable pharmacologic treatments.
Introduction to Clinical Medicine 2 (ICM 2): Two semester course that covers advanced clinical communication skills, humanism and compassion, specialized physical examination techniques, advanced medical ethics, medical socioeconomics and healthcare laws.
SUMMER between MS Year Two and GRAD 1: The United States Medical Licensing Examination (USMLE Step I)
Graduate Research (Typically 3-4 years)
These years are devoted to the course work and original research to fulfill requirements established by individual basic science departments for receiving a PhD in the respective discipline.
Medical School Year 3
Third Year core clerkships run concurrently throughout the academic year beginning in late June and ending the following June. The sequence of clerkships is organized on a track basis.
Core Clerkships:
- Family Medicine (includes AHEC rotation): Six weeks
- Internal Medicine: Eight weeks
- Neurology: Four weeks
- Pediatrics: Six weeks
- Psychiatry: Six weeks
- Obstetrics and Gynecology: Six weeks
- Surgery: Eight weeks
- Clinical Electives and/or Vacation: Four weeks, plus two additional weeks that can be used.
Medical School Year 4
Fourth Year runs concurrently throughout the academic year from July to early May.
Required Clinical Rotations Include:
- Acting Internship (AI): Four weeks
- Advanced Cardiac Life Support (ACLS)
- Critical Care: Two weeks
- Palliative Care: One week
- Clinical Electives: 26- to 28-weeks
- Topics in Clinical Medicine
- Residency Interviews
Clerkships and Rotations are completed at multiple clinic sites, including, but not limited to:
- UofL Health - UofL Hospital
- UofL Health - Jewish Hospital
- Norton Children's Hospital
- Norton Hospital
- VA Medical Center
The United States Medical Licensing Examination (USMLE Step 2)
Cassandra Woolley, PhD
Medical School Year 4
| Graduate Department | Microbiology and Immunology |
|---|---|
| Dissertation Title | "Illuminating under-appreciated mechanisms of receptor regulation in human lymphocytes" |
| PhD Mentor | Dr. Thomas C. Mitchell |
| Undergraduate Degree | BS Biology, 2018 University of Kentucky, Lexington, KY |
Dana Oakes
Medical School Year 3
| Graduate Department | Anatomical Sciences and Neurobiology |
|---|---|
| PhD Mentor | Dr. William Guido |
| Undergraduate Degree | BS Biology and Neuroscience, with minor in Medical Sociology, 2018 Indiana University-Purdue University, Indianapolis, IN |
Daniel Nguyen
Graduate School Year 4
Congratulations, Dan, on receiving a F30 Fellowship Award from the National Institutes of Health, National Heart, Lung and Blood Institute with his Project Title: "Fibroblast TAK1 signaling in cardiac fibrosis".
| Graduate Department | Department of Physiology |
|---|---|
| PhD Mentor | Dr. Bradford G. Hill |
| Undergraduate Degree | BS Applied Exercise Science, 2017 University of New England, Biddeford, ME |
Eric G. Engelbrecht
Medical School Year 3
| Graduate Department | Biochemistry and Molecular Genetics |
|---|---|
| PhD Mentor | Dr. Corey T. Watson |
| Undergraduate Degree | BA Molecular Biology, 2014 Kenyon College, Gambier, OH |
Kara R. Gouwens
Graduate School Year 3
| Graduate Department | Physiology |
|---|---|
| PhD Mentor | Dr. Matthew A. Nystoriak |
| Undergraduate Degree | BS Molecular Biology, 2018 Bethel College, Mishawaka, IN |
Steven T. Nadakal
Graduate School Year 2
| Graduate Department | Biochemistry and Molecular Genetics |
|---|---|
| PhD Mentor | Dr. Melissa L. Smith |
| Intramural Research Fellowship | National Institutes of Health, Department of Health and Human Services, Bethesda, MD, 2018-2021 |
| Graduate Degree | MS Immunology of Infectious Disease, 2018 London School of Hygiene and Tropical Medicine, London, United Kingdom of Great Britain and Northern Ireland |
| Undergraduate Degree | BS Biochemistry, 2017 Baylor University, Waco, TX |
Phillip Harter
Graduate School Year 1
| Fulbright Research Fellowship | Luxembourg Institute of Health, Luxembourg, 2021-2022 |
|---|---|
| Undergraduate Degree | BA Political Science, Minor Biological Sciences, 2021 Purdue University, West Lafayette, IN |
Sidney R. Johnson
Graduate School Year 1
| Graduate Degree | MS Medical Sciences, 2021 University of Kentucky, Lexington, KY |
|---|---|
| Undergraduate Degree | BA Psychology and Theatre, 2017 University of Kentucky, Lexington, KY |
Luke Schroeder
Graduate School Year 1
Megan Bezold
Medical School Year 2
| Undergraduate Degree | BS Neuroscience, minor Public Health, 2023 University of Louisville, Louisville, KY |
|---|
Ilaria Ferrari
Medical School Year 2
| Undergraduate Degree | BS Mechanical Engineering, 2018 Columbia University, New York, NY |
|---|
Idil Tuncali
Medical School Year 2
| Graduate Degree | MS Neuroscience, 2018 University of Massachusetts, Amherst, MA |
|---|---|
| Undergraduate Degree | BS Neuroscience, 2016 University of Massachusetts, Amherst, MA |
Theresa Weis
Medical School Year 2
| Undergraduate Degree | BS Biology, minor Chemistry, 2020 Brandeis University, Waltham, MA |
|---|
Matilyn Shanahan
Medical School Year 1
Whitney Heard
Medical School Year 1
Zoe Hopewell
Medical School Year 1
Liam Scott
Medical School Year 1
Graduates of the program have had outstanding success in matching high quality residencies and transitioning into prominent fellowships.
Evan Tracy, PhD
May 2024
| Residency | Johns Hopkins University, Internal Medicine Combined Physician Scientist Training Program |
|---|---|
| Graduate Department | Physiology |
| Dissertation Title | "Stromal Vascular Fraction Restores Vasodilatory Function By Reversing Mitochondrial Dysfunction and Oxidative Stress In Aging-Induced Coronary Microvascular Disease" |
| PhD Mentor | Dr. Amanda Jo LeBlanc |
| Undergraduate Degree | BS Biology, 2015 Linfield College, McMinnville, OR |
Jordan Noe, MD, PhD
May 2023
| Residency | University of Louisville, Louisville, KY, General Surgery Residency Program, Preliminary |
|---|---|
| Graduate Department | Biochemistry and Molecular Genetics |
| Dissertation Title | "Metabolic-Epigenetic Regulation of Macrophage Polarization" |
| PhD Mentor | Dr. Robert A. Mitchell |
| F30 AWARD | National Institutes of Health, National Cancer Institute, Project Title: "The role of MIF in mitochondrial metabolism and M2-TAM polarization." |
| Undergraduate Degree | BS Biology, minor in Chemistry, 2013 University of Utah, Salt Lake City, UT |
Anne Geller, MD, PhD
May 2023
| Residency | Johns Hopkins University, Baltimore, MD, Urology Residency Program |
|---|---|
| Graduate Department | Microbiology and Immunology |
| Dissertation Title | "Harnessing the Power of Trained Immunity in the Setting of Pancreatic Cancer: A Novel Mechanism of Immune Trafficking and Tumor Control" |
| PhD Mentor | Dr. Jun Yan |
| Intramural Research Fellowship | National Institutes of Health, Department of Rehabilitation Medicine, Bethesda, MD, 2014-2015 |
| Graduate Degree | MS Biomedical Science, 2014 Tufts University, Medford, MA |
| Undergraduate Degree | BS Biomedical Science in Engineering, minor in Engineering Entrepreneurship, 2013 University of Pennsylvania, Philadelphia, PA |
George Wei, MD, PhD
May 2023
| Residency | Emory University, Atlanta, GA, Neurosurgery Residency Training Program |
|---|---|
| Graduate Department | Pharmacology and Toxicology |
| Dissertation Title | "Oligodendrocyte Responses after Spinal Cord Injury" |
| PhD Mentors | Dr. Michal Hetman and Dr. Scott Whittemore |
| Undergraduate Degree | BA Chemistry, 2014 Rutgers University, New Brunswick, NJ |
Desmond Harrell-Stewart, MD, PhD
May 2022
| Residency | University of Maryland, Baltimore, MD, for Internal Medicine Research Pathway |
|---|---|
| Graduate Department | Pharmacology and Toxicology |
| Dissertation Title | "A Novel Role for RASSF1A in the Regulation of Ras Activation" |
| PhD Mentor | Dr. Geoffrey Clark |
| Undergraduate Degree | BS Biochemistry, minor in Biology, 2014 Xavier University of Louisiana, New Orleans, LA |
Samantha Morrissey, MD, PhD
May 2022
| Residency | University of North Carolina, Chapel Hill, NC, for Internal Medicine/Pediatrics Residency Program |
|---|---|
| Graduate Department | Microbiology and Immunology |
| Dissertation Title | "Tumor-derived Exosomes Drive Immunosuppressive Macrophages in a Pre-metastic Niche through NF-kB dependent Glycolytic Metabolic Reprogramming" |
| PhD Mentor | Dr. Jun Yan |
| Undergraduate Degree | BA Biology, minor in Chemistry, 2013 University of Pennsylvania, Philadelphia, PA |
Peter Campbell, MD, PhD
May 2021
| Residency | Baylor College of Medicine, Houston, TX for Child Neurology, receiving one of two spots, with one year of basic neuroscience research |
|---|---|
| Graduate Department | Anatomical Sciences and Neurobiology |
| Dissertation Title | "Development, form and function of intrathalamic circuits" |
| PhD Mentor | Dr. William Guido |
| F30 AWARD | National Institutes of Health, National Eye Institute, Project Title: "Formation and Maturation of Projections Between Thalamic Reticular Nucleus and Dorsal Lateral Geniculate Nucleus" |
| Undergraduate Degree | BA Independent Study in Neuroscience, 2006 Grinnell College, Grinnell, IA |
Douglas Saforo, MD,PhD
May 2021
| Residency | Emory University, Atlanta, GA Pediatrics Residency Training Program |
|---|---|
| Graduate Department | Pharmacology and Toxicology |
| Dissertation Title | "Characterizing the role of extracellular matrix in non-small cell lung carcinoma metastasis using a 3D microenvironment mimetic in-vitro culture system" |
| PhD Mentor | Dr. Leah Siskind and Dr. Levi Beverly |
| Undergraduate Degree | BA Biology, 2013 University of Louisville, Louisville, KY |
Heather Clair, MD, PhD
May 2019
Dr. Clair is a board-certified family medicine physician who joined Flagler Health and is currently associated with Primary Care at Julington Creek in St. Johns, Florida.
| Residency | Orange Park Medical Center, Orange Park, FL, Family Medicine, 2022 |
|---|---|
| Graduate Department | Biochemistry and Molecular Genetics |
| Dissertation Title | "PCB-Associated Steatohepatitis and the Role of Xenobiotic Receptors" |
| PhD Mentor | Dr. Matthew Cave |
| F30 AWARD | National Institutes of Health, National Institute of Environmental Health Sciences, Project Title: "Xenobiotic Receptors Modulate PCB-induced Steatohepatitis" |
| Graduate Degree | MS Animal Science, Biotechnology, 2001 University of Georgia, Athens, GA |
| Undergraduate Degree | BS Animal Science, Dairy Science, 1999 University of Georgia, Athens, GA |
Venkat Ramakrishnan, MD, PhD
May 2018
| Fellowship | Pediatric Urology Fellowship, Boston Children's Hospital, Harvard University Medical School, Boston, MA |
|---|---|
| Residency | Harvard University, Brigham and Women's Hospital, Boston, MA - Urology Residency Program. Dr. Ramakrishnan was selected to be one of the 2022-2023 chief residents and will stay for an additional year to fulfill that role. |
| Graduate Department | Department of Physiology |
| Dissertation Title | "Development of a Vascularized, Induced Pluripotent Stem Cell-Derived Liver-Tissue Mimic for Therapeutic Applications" |
| PhD Mentor | Dr. Nolan Boyd |
| Fulbright Research Fellow | University Hospital, Zurich, Switzerland, 2010-2011 |
| Undergraduate Degree | BS Biology (Cellular Physiology), 2010 University of Louisville, Louisville, KY |
Jessica Mezzanotte Sharpe, MD, PhD
May 2018
| Fellowship | Hematology/Medical Oncology Fellowship, Department of Medicine, Division of Hematology/Oncology. Vanderbilt University, Nashville, TN |
|---|---|
| Residency | The Ohio State University Medical Center, Columbus, OH, Internal Medicine Residency Program. Dr. Sharpe was selected to be one of the 2021-2022 chief residents and stayed for an additional year to fulfill that role. 2018-2022 |
| Graduate Department | Department of Biochemistry and Molecular Genetics |
| Dissertation Title | "The discovery of a novel, Ras-mediated NORE1A/PMLIV tumor suppressor complex" |
| PhD Mentor | Dr. Geoffrey Clark |
| Undergraduate Degree | BA Biology, minors in Chemistry and English, 2010 Vanderbilt University, Nashville, TN |
Huy T. Le, MD, PhD
May 2016
| Faculty Appointment | Assistant Professor, Department of Internal Medicine, Division of Hospital Medicine at the University of Nebraska Medical Center, Omaha, NE |
|---|---|
| Residency | University of Nebraska Medical Center, Omaha, Nebraska - Internal Medicine Pediatrics Residency Program, 2020 |
| Graduate Department | Department of Biochemistry and Molecular Biology |
| Dissertation Title | "Biophysical and Computational Investigation into G-Quadruplex Structural Polymorphism and Interaction with Small Molecules." |
| PhD Mentor | Dr. John Trent |
| Undergraduate Degree | BS Chemistry, minor in Biology; University of Louisville, Louisville, KY |
Michael J. Zhang, MD, PhD
May 2016
| Fellowship | University of Minnesota, Minneapolis, MN - Physician Scientist Training Program Fellowship Dr. Zhang was awarded a NIH F32 training grant and will spend 3 years in the lab before finishing his fellowship in 2024. He was also awarded an American Heart Association Postdoctoral Fellowship Grant in 2021. |
|---|---|
| Residency | University of Minnesota, Minneapolis, MN - Physician Scientist Training Program - two years of clinical training (short track) as an Internal Medicine resident. |
| Graduate Department | Department of Physiology and Biophysics |
| Dissertation Title | "Inflammation-resolving Lipid Mediators Promote Revascularization to Enhance Wound Healing" |
| PhD Mentors | Dr. Aruni Bhatnagar and Dr. Matthew Spite |
| Undergraduate Degree | BS Microbiology, Immunology and Molecular Genetics, minor in Global Studies University of California, Los Angeles, CA |
Alex Belshoff, MD, PhD
May 2015
Dr. Belshoff joined Aloha Urology in Honolulu, Hawaii in August 2021.
| Residency | Loyola University Medical Center, Chicago, IL - Urology Residency Program, 2021 |
|---|---|
| Graduate Department | Department of Pharmacology and Toxicology |
| Dissertation Title | "Probing the Anti-cancer Mechanism of Selenite: a Metabolic Approach" |
| PhD Mentor | Dr. Teresa Fan |
| Undergraduate Degree | BS Chemistry, University of Louisville, Louisville, KY |
Alden Klarer, MD, PhD
May 2015
| Faculty Appointment | Assistant Professor, University of Louisville, Department of Radiation Oncology, Louisville, KY Dr. Klarer specializes in the treatment of breast malignancies and is involved in a number of investigator-initiated and cooperative group clinical trials. Her treatment expertise and clinical interests includes intensity modulated radiation therapy, cardiac-sparing radiation therapy, stereotactic body radiotherapy and radiobiology |
|---|---|
| Residency | University of Louisville, Department of Radiation Oncology, Radiation Oncology Residency, 2020 University of Louisville, Louisville, KY for preliminary year in Internal Medicine, 2016 |
| Graduate Department | Department of Biochemistry and Molecular Biology |
| Dissertation Title | "6-Phosphofructo-2-Kinase Inhibition Induces Autophagy as a Survival Mechanism" |
| PhD Mentors | Dr. Steven Ellis and Dr. Sucheta Telang |
| F30 AWARD | Grant award from the National Institute of Environmental Health Sciences, Project title: "Y-family DNA Polymerases and Cellular Responses to Benzo[a]pyrene" |
| Undergraduate Degree | BS Biology, University of Kentucky, Lexington, KY, 2004 |
Francesca Rinaldo, MD, PhD
May 2015
Dr. Rinaldo is an Affiliated Scholar at the Stanford Clinical Excellence Research Center (CERC) and Senior Vice President of Clinical Product and Innovation at Sharecare. In her role at Sharecare, she is the Chief Medical Officer of CareLinx.
| Fellowship | Stanford University, Stanford, CA - Clinical Excellence Research Center (CERC) Fellowship 2017-2018 Dr. Rinaldo put her surgical residency on hold to accept a year long fellowship as a Design Fellow. CERC is a value-based healthcare "think-tank" with a focus on using scientific evidence to design less costly approaches and innovating models of healthcare delivery to improve the quality of care and safely and significantly reduce costs. Upon completing the fellowship, she remained at CERC as an Associate Director for the Healthcare Design Fellowship and served as a Clinical Lead for the Stanford CERC Partnership for AI Assisted Care (PAC). At PAC, she co-led a team of computer scientists and clinicians in a study examining the application of computer vision deep-learning algorithms to detect patterns of mobility in critically ill patients and how these patterns drive post-ICU clinical outcomes. This work was published in Nature Digital Medicine in March 2019. |
|---|---|
| Residency | Stanford University, Division of General Surgery, Stanford, CA - Accelerated Surgeon Scientist Track Dr. Rinaldo finished her first two years of surgical residency and was voted the Consult Resident of the Year by her surgical attendings at the end of year two. |
| Graduate Department | Department of Biochemistry and Molecular Biology |
| Dissertation Title | "Investigating the Mechanism of Action of XB05: A Novel Small Molecule with Anticancer Properties" |
| PhD Mentor | Dr. Paula Bates |
| Undergraduate Degree | BA Pre-Med, Public Policy Studies, University of Chicago, Chicago, IL |
Alan Brooks, MD, PhD
May 2014
Dr. Brooks is currently associated with Norton Healthcare, Norton Heart and Vascular Institute, Cardiology Program, Louisville, KY.
| Fellowship | University of Rochester, Rochester, New York Cardiovascular Diseases: Fellowship in Research, 2021; Fellowship in Cardiovascular Disease, 2020 |
|---|---|
| Residency | University of Rochester, Strong Memorial Hospital, Rochester, NY - Physician Scientist Training Program, Internal Medicine Research Path, 2016 |
| Graduate Department | Department of Biochemistry and Molecular Biology |
| Dissertation Title | "Unfolded Protein Response Signaling in Myocardial Response to Ischemia" |
| PhD Mentor | Dr. Aruni Bhatnagar |
| Graduate Degree | ME Chemical Engineering, University of Louisville, Louisville, KY |
| Undergraduate Degree | BS Chemical Engineering, University of Louisville, Louisville, KY, 2003 |
Lacey Jo Gunn, MD, PhD
May 2014
| Faculty Appointment | Assistant Professor, University of Louisville, Department of Family and Geriatric Medicine, Louisville, KY |
|---|---|
| Residency | University of Louisville, Louisville, KY, Department of Family and Geriatric Medicine, 2017 |
| Graduate Department | Department of Microbiology and Immunology |
| Dissertation Title | "Effect of Complement C5a in Tumor Progression and the Local Tumor Microenvironment" |
| PhD Mentor | Dr. Jun Yan |
| Undergraduate Degree | BA Biology and Chemistry, Earlham College, Richmond, IN |
Courtney Jetun Mitchell, MD, PhD
May 2014
| Faculty Appointment | Assistant Professor, University of Mississippi Medical Center, School of Medicine, Department of Obstetrics and Gynecology, Maternal and Fetal Medicine Fellowship Faculty, Jackson, MS. Dr. Mitchell's research interests include: hypertensive disorders in pregnancy, maternal mortality and morbidity, barriers to care in resource poor areas, OB quality improvement. |
|---|---|
| Fellowship | Maternal Fetal Medicine Fellowship, Duke University School of Medicine, Department of Obstetrics and Gynecology, 2021 |
| Residency | University of Alabama Medical Center, Birmingham, AL Department of Obstetrics and Gynecology Residency, 2018 |
| Graduate Department | Department of Microbiology and Immunology |
| Dissertation Title | "IL12 Induced Modifications of Tumor Infiltrating Macrophages" |
| PhD Mentors | Dr. Robert Stout and Dr. Jill Suttles |
| F31 AWARD | Grant award from the National Cancer Institute, Project title: "Functional Conversion of Tumor Associated Macrophages via Cytokine Therapy" |
| Undergraduate Degree | BS Biology, Clark Atlanta University, Atlanta, GA, 2007 |
Adrianna Henson Masters, MD, PhD
May 2014
| Faculty Appointment | Assistant Professor, University of Louisville, Department of Radiation Oncology, Louisville, KY |
|---|---|
| Residency | Wake Forest Baptist Medical Center, Winston-Salem, NC for Radiation Oncology University of Louisville, Louisville, KY for Preliminary Year Internal Medicine, 2015 |
| Graduate Department | Department of Biochemistry and Molecular Biology |
| Dissertation Title | "Molecular Bases of Inherited Bone Marrow Failures: Shwachman-Diamond Syndrome and Diamond Blackfan Anemia" |
| PhD Mentor | Dr. Steven Ellis |
| Undergraduate Degree | BS Chemistry, minor in Biology, Wake Forest University, Winston-Salem, NC, 2005 |
Carlo Renato G. Bartoli, MD, PhD
May 2013
| Faculty Appointment | Assistant Professor, Geisinger Commonwealth School of Medicine, Geisinger Medical Center, Cardiothoracic Surgery, Danville, PA Dr. Bartoli is a cardiac surgeon-scientist with special interests in surgical management of congenital and acquired heart disease, development and clinical implantation of mechanical circulatory support devices, von Willebrand factor biology and angiogenesis. His appointment is 60% clinical and 40% research. In parallel with his clinical practice, Dr. Bartoli manages an internationally-recognized translational biophysics laboratory that is independently funded with support from the NIH, multiple congenital cardiology foundations and industry partners. Ongoing projects examine mechanical circulatory support in the fetus to the adult and include left ventricular assist device (LVAD) design and development, mechanisms of LVAD-associated bleeding and thrombosis, novel LVAD therapies, abnormal angiogenesis in children with single ventricle disease and cardiovascular biology of the placenta. |
|---|---|
| Faculty Appointment | Instructor of Surgery, University of Pennsylvania, Philadelphia, PA Division of Cardiothoracic Surgery, with a translational biophysics laboratory Awarded the 2020 American College of Cardiology Doug Zipes Distinguished Young Scientist Award for "contributions to cardiovascular medicine and significant body of research." |
| Fellowship | Pediatric Cardiothoracic Surgery, Children's Hospital of Philadelphia, PA, June 2020 |
| Residency | Integrated Cardiothoracic Surgery Residency, University of Pennsylvania, Philadelphia, PA, June 2019. Dr. Bartoli was the Chief Resident of the Cardiovascular Surgery service. He received the Young Author Achievement Award from the American College of Cardiology, in recognition of outstanding research published in the Journal of the American College of Cardiology: Heart Failure. |
| Graduate Department | Department of Physiology and Biophysics |
| Dissertation Title | "Partial vs Full Support of the Heart with a Continuous-Flow Left Ventricular Assist Device: Implications for Myocardial Recovery" |
| PhD Mentors | Dr. Steven Koenig and Dr. Sumanth Prabhu |
| Graduate Degree | MA Biotechnology, Harvard University, Cambridge, MA |
| Undergraduate Degree | BS Biology (Physiology), Cornell University, Ithaca, NY, 2004 |
Toros A. Dincman, MD, PhD
May 2013
| Faculty Appointment | Assistant Professor, Medical University of South Carolina, Charleston, SC, Department of Medicine, Division of Hematology and Oncology Dr. Dincman was also appointed as a South Carolina Translational Research (SCTR) Institute KL2 Scholar which provides protected research time/salary support and research funds. |
|---|---|
| Fellowship and Residency | Medical University of South Carolina (MUSC), Charleston, SC ABIM subspecialty research pathway in Hematology/Oncology which included appointment as a T32 post-doctoral scholar in Integrative Training in Oncogenic Signaling (ITOS), 2020 |
| Graduate Department | Department of Anatomical Sciences and Neurobiology |
| Dissertation Title | "Histone deacetylase inhibition-mediated cytotoxicity oligodendrocytes" |
| PhD Mentor | Dr. Scott Whittemore |
| Undergraduate Degree | BA Biological Sciences, University of Chicago, Chicago, IL, 2004 |
Enid Choi, MD, PhD
May 2012
| Faculty Appointment | Assistant Professor, University of Michigan, Department of Radiation Oncology, Ann Arbor, MI at VA Ann Arbor Healthcare System |
|---|---|
| Residency | University of Michigan, Department of Radiation Oncology, Ann Arbor, MI (2015-2017) University of Maryland, Baltimore, MD - Radiation Oncology (2013-2015) University of Louisville, Louisville, KY - Preliminary year in Internal Medicine, 2013 |
| Graduate Department | Department of Biochemistry and Molecular Biology |
| Dissertation Title | "Tumor-targeting aptamers for the treatment of prostate cancer" |
| PhD Mentor | Dr. Paula Bates |
| Undergraduate Degree | BS Chemical Engineering, Massachusetts Institute of Technology, Cambridge, MA, 2000 |
Janelle Fassbender Adeniran MD, PhD
May 2012
Dr. Adeniran provides vitreoretinal services at Bennett and Bloom Eye Centers, Louisville, KY and maintains a strong interest in clinical research and teaching. She is a Gratis Faculty Member at the University of Louisville, Department of Ophthalmology and Visual Sciences, Louisville, KY.
| Fellowship | University of Louisville, Louisville, KY - Department of Ophthalmology and Visual Science Fellow in Vitreoretinal Diseases and Surgery, 2018 |
|---|---|
| Residency | University of Louisville, Louisville, KY in Ophthalmology, 2016 Good Samaritan Hospital, Cincinnati, OH - Preliminary year in General Surgery, 2013 |
| Graduate Department | Department of Anatomical Sciences and Neurobiology |
| Dissertation Title | "Improving Functional Recovery Following Spinal Cord Injury by Therapeutically Targeting the Vasculature" |
| PhD Mentor | Dr. Scott Whittemore |
| F30 AWARD | Grant award from the National Institute of Neurological Disorders and Stroke, Project title: "Functional Consequences of Notch-mediated Angiogenesis following SCI" |
| Undergraduate Degree | BS Neuroscience, Davidson College, Davidson, NC, 2005 |
Patrick Andrew Holt, MD, PhD
May 2012
Dr. Holt is the Chief of the Department of Surgery, TidalHealth Peninsula Regional. He is an Orthopaedic Surgeon specializing in Hand, Wrist and Elbow nerve decompression surgery, trauma and complex reconstruction at Peninsula Orthopaedic Associates. He is one of only a few fellowship-trained hand surgeons on the Lower Eastern Shore.
| Fellowship | Duke University, Durham, NC - Hand and Microvascular Surgery Program, 2018 |
|---|---|
| Residency | University of Utah, Salt Lake City, UT in Orthopaedic Surgery, 2017 |
| Graduate Department | Department of Biochemistry and Molecular Biology |
| Dissertation Title | "Determination of In Silico Rules for Predicting Small Molecule Binding Behavior to Nucleic Acids In Vitro" |
| PhD Mentor | Dr. John Trent |
| Graduate Degree | MS Biotechnology, Johns Hopkins University, Baltimore, MD |
| Undergraduate Degree | BS Biomedical Engineering, Johns Hopkins University, Baltimore, MD |
Justin R. Kingery, MD, PhD
May 2012
| Faculty appointment | Associate Professor of Medicine and Hospitalist, University of Louisville, Department of Medicine, Division of General Internal Medicine and Medical Education. Louisville, KY. |
|---|---|
| Faculty appointment | Assistant Professor in Medicine and Hospitalist, Joan and Sanford I Weill Department of Medicine, Weill Cornell Medical College, New York, NY. Medical Director of the 5W Stepdown Unit at NY Presbyterian Hospital and course instructor at Weill Cornell Medical College. Director, Global Ultrasound at Weill Cornell Medical College and Center for Global Health. Lecturer in Medicine at Weill Bugando College of Health Sciences, Mwanza, Tanzania and the Global Health Center in Port-au-Prince, Haiti. |
| Fellowship | Global Health Research Fellow at the Weill Cornell Medical College/New York Presbyterian Hospital and the Weill Bugando College of Health Sciences, Mwanza,Tanzania |
| Residency | University of Louisville, Louisville, KY in Internal Medicine, 2015 |
| Graduate Department | Department of Physiology and Biophysics |
| Dissertation Title | "Inflammatory Cell-Derived Inducible Nitric Oxide Synthase Exacerbates Post-Inarction Left Ventricula Remodeling" |
| PhD Mentor | Dr. Sumanth Prabhu |
| Undergraduate Degree | BS Chemistry, Biochemistry, Marshall University, Huntington, WV |
Jason B. Meier, MD, PhD
May 2012
Dr. Meier is a Hematologist/Oncologist at the Cancer Center at Cookeville Regional Medical Center, Cookeville, TN
| Fellowship | Vanderbilt University, Nashville, TN, Hematology/Oncology Fellowship Program |
|---|---|
| Residency | Indiana University (IUPUI) in Internal Medicine |
| Graduate Department | Department of Biochemistry and Molecular Biology |
| Dissertation Title | "Structure Based Drug Design for CXCR4" |
| PhD Mentor | Dr. John Trent |
| Undergraduate Degree | BS Biology, Chemistry, University of Louisville, Louisville, KY, 2004 |
Maria E. Barnes-Davis, MD, PhD
May 2011
| Faculty Appointment | Associate Professor, University of Cincinnati, Department of Pediatrics in the Perinatal Institute at Cincinnati Children's Hospital Medical Center and Neuroscience Graduate Program at UC College of Medicine. Dr. Barnes-Davis is an Attending Neonatologist in the Division of Neonatology and Pulmonary Biology and has funding via a K-12 Award through Eunice Kennedy Shriver National Institute of Child Health and Human Development. Her research focuses on Functional neuroimaging (fMRI, MEG, EEG); neurodevelopmental outcomes of extremely preterm children; language; sleep and learning disorders. |
|---|---|
| Clinical Fellowship | Neonatal and Pulmonary Biology, Cincinnati Children’s Hospital Medical Center, Cincinnati, OH |
| Residency | Icahn School of Medicine at The Mount Sinai Medical Center, New York, NY in Integrated Pediatrics/Research Track. |
| Graduate Department | Department of Psychological and Brain Sciences |
| Dissertation Title | "Pre-and Post-operative Differences in Brain and Sleep-related Events in Children" |
| PhD Mentors | Dr. David Gozal and Dr. Dennis Molfese |
| F30 AWARD | Grant award from the National Institute of Mental Health, Project title: "Pre- and Post-operative Differences in Brain and Sleep-related Events in Children with OSA" |
| Undergraduate Degree | BA Philosophy, Brown University, Providence, RI, 2003 |
W. Lee Titsworth, MD, PhD, MPH
May 2010
Neurosurgeon Lee Titsworth, MD, PhD, joined the Baptist Health Medical Group Neurosurgery in Paducah, KY as the only fellowship-trained pediatric neurosurgeon in the region. Dr. Titsworth provides surgical treatment for conditions of the brain and spine, with emphasis in adult and adolescent scoliosis surgery and endoscopic brain surgery.
| Fellowship | Johns Hopkins University, Pediatric Neurosurgery Fellowship, 2017 |
|---|---|
| Residency | University of Florida/Shands Medical Center, Gainesville, FL in the Neurosurgical Residency Program, 2016 |
| MPH | Clinical Effectiveness, Harvard University, School of Public Health, Fall 2015 |
| Graduate Department | Department of Anatomical Sciences and Neurobiology |
| Dissertation Title | "A Possible Role for sPLA2 in Oligodendrocyte Death and Spinal Cord Injury" |
| PhD Mentor | Dr.Xiao-Ming Xu |
| F31 AWARD | Grant award from the National Institute of Neurological Disorders and Stroke, Project title: "Role of sPLA2 in Post Spinal Cord Injury Demyelination" |
| Undergraduate Degree | BA Psychology, Samford University, Birmingham, AL, 2001 |
Christina B. Clark, MD, PhD
May 2010
| Faculty Appointment | Assistant Professor, University of Louisville, Department of Radiology, Breast Imaging, Louisville, KY |
|---|---|
| Fellowship | University of Rochester, Rochester, New York, Breast Imaging Fellowship, 2016-2017, where she had the opportunity to work on a clinical research project focused on the USA's only FDA-approved cone beam breast CT scanner at UR's Breast Imaging Center. |
| Residency | Geisinger Medical Center, Danville, PA in Diagnostic Radiology Residency Program, 2016 University of Cincinnati, Cincinnati, OH - Preliminary year in Internal Medicine, 2011 |
| Graduate Department | Department of Pharmacology and Toxicology |
| Dissertation Title | "Role of Hsp90 Interactions in PC-12 Neuronal Cell Survival to Hypoxia" |
| F30 AWARD | Grant award from the National Institute of Neurological Disorders and Stroke, Project title: "Role of Hsp90 Interactions in PC-12 Survival to Hypoxia" |
| PhD Mentor | Dr. Evelyne Gozal |
| Undergraduate Degree | BA Molecular Biology & Biochemistry, German Literature, Washington University, St. Louis, MO |
B. Frazier Taylor, MD, PhD
May 2009
Dr. Taylor is a Radiation Oncologist with Northeastern Radiation Oncology, PLLC. He provides services at Saratoga Hospital's Mollie Wilmot Radiation Oncology Center in Saratoga Springs, NY; the C. R. Wood Cancer Center in Glens Falls, NY.; and Elllis Hospital, Schenectady, NY.
| Residency | Yale University School of Medicine, Department of Therapeutic Radiology Radiation Oncology Training Program, 2014 |
|---|---|
| Graduate Department | Department of Pharmacology and Toxicology |
| Dissertation Title | "Arsenite Inhibition of Mitotic Progression" |
| PhD Mentor | Dr. J. Christopher States |
| Undergraduate Degree | BS Biochemistry, minor in Studio Art, Denison University, Granville, OH |
James Massey, MD, PhD
May 2008
In August 2014, Dr. Massey returned to his hometown of Nashville, TN and joined Radiology Alliance, P.C., specializing in diagnostic and pediatric radiology. Radiology Alliance, P.C., is a provider of inpatient and outpatient radiology services at TriStar Health System's nine Middle Tennessee Hospitals.
| Fellowship | Indiana University, (IUPUI) Department of Radiology and Imaging Sciences, Pediatric Radiology |
|---|---|
| Residency | Cleveland Clinic Imaging Institute, Cleveland, OH, Diagnostic Radiology Residency Program Transitional year - Akron General Medical Center, Akron, OH |
| Graduate Department | Department of Anatomical Sciences and Neurobiology |
| PhD Mentor | Dr. Stephen Onifer |
| Graduate Degree | MA Perception and Sensory Physiology, University of Louisville, Louisville, KY |
| Undergraduate Degree | BS Biology, minor in Chemistry, David Lipscomb University, Nashville, TN |
Kelby B. Napier, MD, PhD
May 2008
| Faculty Appointment | Assistant Professor, Washington University School of Medicine, Department of Radiology, Division of Musculoskeletal Radiology, St. Louis, MO Dr. Napier specializes in musculoskeletal imaging and MRI, musculoskeletal CT, musculoskeletal ultrasound, image-guided procedures, sports medicine, arthritis and bone and soft tissue tumors. |
|---|---|
| Fellowship | Washington University School of Medicine, St. Louis, MO, Mallinckrodt Institute of Radiology, Musculoskeletal Imaging and Interventions Program |
| Residency | Washington University School of Medicine, St. Louis, MO, Mallinckrodt Institute of Radiology, Diagnostic Radiology Training Program, June 2013 |
| Graduate Department | Department of Biochemistry and Molecular Biology |
| PhD Mentor | Dr. John Trent |
| Undergraduate Degree | BS Chemistry, Biology, Eastern Kentucky University, Richmond, KY |
Stephen R. Reeves, MD, PhD
May 2008
| Faculty Appointment | Assistant Professor of Pediatrics, University of Washington, Department of Pediatrics, Division of Pulmonary and Sleep Medicine, Seattle, WA Attending physician at Seattle Children's Hospital in the Division of Pulmonary and Sleep Medicine. Dr. Reeves continues to do basic science and translational research in the realm of asthma and airway biology at Seattle Children's Research Institute, Center for Immunity and Immunotherapies. |
|---|---|
| Fellowship | University of Washington, Seattle, WA, Seattle Children's Hospital, Pediatric Pulmonology/Respiratory Medicine, June 2014 |
| Residency | University of California, San Diego, CA in General Pediatrics |
| Internship | University of California, San Diego, CA in General Pediatrics, 2009 |
| Graduate Department | Department of Pharmacology and Toxicology |
| Dissertation Title | "Chronic Intermittent Hypoxia and Respiratory Plasticity" |
| F30 AWARD | Grant award from the National Institute of Neurological Disorders and Stroke, Project title: "Postnasal Intermittent Hypoxia and Respiration: Potential Role of Protein Kinase C." |
| PhD Mentor | Dr. David Gozal |
| Undergraduate Degree | BS Biological and Environmental Sciences, Morehead State University, Morehead, KY, 2001 |
Daniel Allendorf, MD, PhD
May 2007
Dr. Allendorf joined Alabama Oncology in August 2013, the largest, private, community-based provider of cancer care services in Alabama. Their network is comprised of nine clinics strategically located throughout central Alabama. He is located at the Shelby Cancer Care Center, Alabaster, AL.
| Fellowship | University of Alabama, Birmingham, AL in Division of Hematology and Medical Oncology, 2013 He received The J. Claude Bennett Award for Excellence in research by an Associate Fellow, 2013. |
|---|---|
| Residency | University of Alabama, Birmingham, AL Internal Medicine Residency Program, Board Certification in Internal Medicine, 2010 He received the William E. Dismukes Award of Excellence for Outstanding Senior Resident, 2010. |
| Graduate Department | Department of Microbiology and Immunology |
| Dissertation Title | "Cellular and Molecular Mechanisms of Action of Beta-Glucans for Tumor Immunotherapy in Syngeneic and Translational Animal Models" |
| PhD Mentors | Dr. Gordon Ross and Dr. Jun Yan |
| Undergraduate Degree | BA Chemistry, Bellarmine University, Louisville, KY |
Anwar Husain, MD, PhD
May 2007
Dr. Husain is a Toxicologist for the US Food and Drug Administration, Center for Tobacco Products (CTP), Office of Science, Division of Nonclinical Science, Silver Spring, MD.
| Postdoctoral Fellowship | Johns Hopkins Medicine, Sidney Kimmel Comprehensive Cancer Center, Bloomberg Kimmel Institute for Cancer Immunotherapy, Department of Oncology - Division of Immunology/Hematopoesis. Developed machine learning based workflows to assist translational research efforts in immuno-oncology research, drug development and discovery and digital pathology whole slide image analysis. |
|---|---|
| Postdoctoral Fellowship | Georgetown University Lombardi Comprehensive Cancer Center Metabolomic and Toxicology Research Scientist under mentorship of Peter G. Shields, M.D., working on metabolomics studies related to cigarette smoke toxicology and folate metabolomics related to breast cancer risk. |
| Graduate Department | Department of Pharmacology and Toxicology |
| Dissertation Title | "Regulatory Control Regions of Human Arylamine N-acetyletransferase 1 and 2: Implications for Genetic Implications to Breast Cancer" |
| F30 AWARD | Grant award from the National Institute of Environmental Health Sciences, Project title: "Genetic Polymorphisms in 5'-UTR of Human NAT1 and NAT 2" |
| PhD Mentor | Dr. David Hein |
| Graduate Degree | MS Computer Science, Hood College, Frederick, MD 2018 MS Physiology and Biophysics, Georgetown University, Washington, DC, 2000 |
| Undergraduate Degree | BS Computer Science, University of Maryland, College Park, MD, 2016 BS Biochemistry, University of Maryland, College Park, MD, 1998 |
Jason Talbott, MD, PhD
May 2007
| Faculty appointment | Associate Professor, Neuroradiology section, Department of Radiology and Biomedical Imaging, University of California, San Francisco and Zuckerberg San Francisco General Hospital (ZSFG). Dr. Talbott is the Director of Clinical Operations at ZSFG. Dr. Talbott also serves as the Associate Residency Program Director, Diagnostic Radiology Residency Program. Research interests include advanced spinal cord and peripheral nerve MR imaging techniques utilizing experimental animal models. |
|---|---|
| Fellowship | University of California, San Francisco, CA, Neuroradiology, 2013 |
| Residency | University of California, San Francisco, CA, Diagnostic Radiology Residency, 2012 In 2012 he received the Alexander Margulis Outstanding Resident Research Award and also served as a Chief Resident. In 2010-2011 he was selected to participate in research training in Biomedical Imaging sponsored by a T32 Training Grant from the National Institute of Biomedical Imaging and Bioengineering. At the 8th Annual Imaging Research Symposium in August 2011, Dr. Talbott received the award for Best Presentation for "Diffusion Weighted-MRI of White Matter Injury Produced by Ethidium Bromide and Cervical Spinal Cord Contusion Injuries in the Rat". |
| Internship | University of California, California Pacific Medical Center, San Francisco, CA, preliminary year in Internal Medicine |
| Graduate Department | Department of Anatomical Sciences and Neurobiology |
| Dissertation Title | "Cellular and Molecular Mechanisms Regulating Remyelination in the Adult Rat Spinal Chord" |
| PhD Mentor | Dr. Scott Whittemore |
| Undergraduate Degree | BS Biology, Stanford University, Stanford, CA, 2000 |
David N. Loy, MD, PhD
Dec 2002
| Faculty appointments | Clinical Instructor in Neuroradiology, Washington University at Mallinckrodt Institute of Radiology, St. Louis, MO as of March 2024. He had been an Assistant Professor, University of Virginia School of Medicine, Charlottesville, VA, Department of Radiology and Medical Imaging, Division of Diagnostic and Interventional Neuroradiology and was the Co-Medical Director of the Comprehensive Stroke Center at Bon Secours St. Mary's Hospital, Richmond, VA Dr. Loy had previously been on faculty at Washington University, Mallinckrodt Institute of Radiology, St. Louis, MO, in Division of Neuroradiology/Interventional Radiology, 2010-2011. |
|---|---|
| Fellowship | Washington University, St. Louis, MO, Mallinckrodt Institute of Radiology, Endosurgical Neuroradiology, 2010 Two year fellowship that included one year of specialized diagnostic neuroradiology followed by a year of interventional training involving endovascular treatment of brain aneurysms and advanced interventional spine procedures. |
| Residency | Washington University, St. Louis, MO, Mallinckrodt Institute of Radiology, Barnes-Jewish Hospital, Diagnostic Radiology Training Program, 2008. The residency tract allowed him to spend 1 full year in the lab, working with Dr. Victor Song in the Biomedical MR laboratory, focusing on MR diffusion tensor imaging in spinal cord injury. Additionally, he started a collaborative effort between WashU and the UofL Dept. of Neurosurgery in 2006. |
| Internship | University of Louisville, Louisville, KY, in General Surgery, 2004 |
| Post-doctoral Research Fellowship | University of Louisville, Louisville, KY, Department of Anatomical Sciences and Neurobiology - mentored by Dr. Scott Whittemore and Dr. Christopher Shields |
| Graduate Department | Department of Anatomical Sciences and Neurobiology |
| Dissertation Title | "Locomotor Pathways and Microvascular Processes Following Spinal Cord Injury" |
| PhD Mentor | Dr. Scott Whittemore |
| Undergraduate Degree | BA Biology, University of Louisville, Louisville, KY |
Publications produced from University of Louisville work by current MD-PhD Students and Recent Graduates
Kritika Bajaj, Sarah A. Andres, Dillon T. Hofsommer, Aidan F. Greene, Oleksandr Heitsoi, Mark S. Mashuta, Theresa Weis, Levi J. Beverly, Paula J. Bates, Robert M. Buchanan, Craig A. Grapperhaus. Physical structure of constitutional isomers influences antiproliferation activity of thiosemicarbazone-alkyl thiocarbamate copper complexes. Journal of Inorganic Biochemistry, 246 (2023). https://doi.org/10.1016/j.jinorgbio.2023.112288
Eric Engelbrecht, Oscar L. Rodriguez, Kaitlyn Shields, Steven Schultze, David Tieri, Uddalok Jana, Gur Yaari, William D. Lees, Melissa L. Smith & Corey T. Watson: Resolving haplotype variation and complex genetic architecture in the human immunoglobulin kappa chain locus in individuals of diverse ancestry. doi:10.1038/s41435-024-00279-2
Brown TC, Crouse EC, Attaway CA, Oakes DK, Minton SW, Borghuis BG , McGee AW. Microglia are dispensable for experience-dependent refinement of visual circuitry. bioRxiv. 2023 Oct 17:2023.10.17.562708. Preprint. doi:10.1101/2023.10.17.562708
Gao Y, Wei GZ, Forston MD, Rood B, Hodges ER, Burke D, Addres K, Morehouse J, Armstrong Cm Glover C, Slomnicki LP, Ding J, Chariker JH, Rouchka EC, Saraswat Ohri S, Whittemore SR, Heman M. Opposite modulation of functional recovery following contusive spinal cord injury in mice with oligodendrocyte-selective deletions of Atf4 and Chop/Ddit3. Sci Rep. 2023 Jun 6;13(1):9193. doi: 10.1038/s41598-023-36258-2.
Keith Rj, Hom RH, Amraotkar AR, Bezold MM, Brick JM, Bushau-Sprinkle AM, Hamorsky KT, Kitterman KT, Palmer KE, Smith T, Yeager R, Bhatnagar A. Stratified Simple Random Sampling Versus Volunteer Community-Wide Sampling for Estimates of COVID-19 Prevalence. Am J Public Health. 2023 Jul;113(7). Epub 2023 May 18. doi: 10.2105/AJPH.2023.307303
Wang Y, Geller AE, Yan J. Spatial TIME landscape and its prognostic value in the lung and brain tumor: location matters. Signal Transduct Target Ther. 2023 May 8;8(1):192. doi: 10.1038/s41392-023-01473-w.
Woeste MR, Shrestha R, Geller AE, Li S, Montoya-Durango D, Ding C, Hu X, Li H, Puckett A, Mitchell RA, Hayat T, Tan M, Li Y, McMasters KM , Martin RCG, Yan J. Irreversible electroporation augments beta-glucan induced trained innate immunity for the treatment of pancreatic ductal adenocarcinoma. J ImmunotherCancer. 2023 Apr;11(4):e006221. doi: 10.1136/jitc-2022-006221
Ford EE, Tieri D, Rodriguez OL, Fancoeur NJ, Sota J, Kos JT, Peres A, Gibson WS, Silver CA, Deikus G, Hudson E, Woolley CR, Beckmann N, Charney A, Mitchell TC, Yaari G, Sebra RP, Watson CT, Smith ML. FLAIRR-Seq: A Method for Single-Molecule Resolution of Near Full-Length Antibody H Chain Repertoires. J Immunol.. 2023 May 15;210(10):1607-1619. doi: 10.4049/jimmunol.2200825.
Noe JT, Ding C, Geller AE, Rendon BE, Yan J, Mitchell RA. A Tumor-admixture Model to Interrogate Immune Cell-dependent Tumorigenesis. Bio Protoc. 2023 Mar 5;13(5):e4630. eCollection 2023 Mar 5. doi: 10.21769/BioProtoc.4630
Engelbrecht E, Kooistra T, Knipe RS. The Vasculature in Pulmonary Fibrosis. Cur Tissue Microenviron Rep. 2022 Dec;3(4):83-97. Epub 2022 Jul 13. doi:10.1007/s43152-022-00040-9. (Available 2023-12-01)
Ding C, Shrestha R, Zhu X, Geller AE, Wu S, Woeste MR, Li W, Wang H, Yuan F, Xu R, Chariker JH, Hu X, Li H, Tieri D, Zhang HG , Rouchka EC, Mitchell R, Siskind LJ, Zhang X, Xu XG, McMasters KM, Yu Y, Yan J. Inducing trained immunity in pro-metastatic macrophages to control tumor metastasis. Nat Immunol. 2023 Feb;24(2):239-254. Epub 2023 Jan 5. doi: 10.1038/s41590-022-01288-8
Whittemore SR, Saraswat Ohri S, Forston MD, Wei GZ, Heman M. The Proteostasis Network: A Global Therapeutic Target for Neuroprotection after Spinal Cord Injury. Cells. 2022 Oct 22;11(21):3339. doi: 10.3390/cells11213339
Wang Y, Geller AE, Yan J. Unexpected Breg-NK crosstalk in STING agonist therapy. Cell Mol Immunol. 2022 Dec;19(12):1330-1332. Epub 2022 Nov 10. doi:1038/s41423-022-00952-4
Woolley CR, Chariker JH, Rouchka EC, Ford EE, Hudson EA, Waigel SJ, Smith ML, Mitchell TC. Reference long-read isoform-aware transcriptomes of 4 human peripheral blood lumphocyte subsets. G3 (Bethesda). 2022 Nov 4:12(11):jkac253. doi: 10.1093/g3journa/jkac253.
Rowe G, Heng DS, Beare JE, Hodges NA, Tracy EP, Murfee WL, Le Blan AJ. Enhanced beta-1 adrenergic receptor responsiveness in coronary arterioles following intravenous stromal vascular fraction therapy in aged rats. J Vasc Res. 2022;59(6):343-357. Epub 2022 Sep 8. doi: 10.1159/000526002.
Tracy EP, Nair R, Roe G, Beare JE, Beyer A, LeBlanc AJ. Adipose Stromal Vascular Fraction Reverses Mitochondrial Dysfunction and Hyperfission in Aging-Induced Coronary Microvascular Disease. Am J Physiol Heart Circ Physiol. 2022 Aug 26. doi: 10.1152/ajpheart.00311.2022.
Tracy EP, Dukes M, Rowe G, Beare JE, Nair R, LeBlanc AJ. Stromal Vascular Fraction Restores Vasodilatory Function by Reducing Oxidative Stress in Aging-Induced Coronary Microvascular Disease. Antioxid Redox Signal. 2022 Aug 11. Epub 2022 Jul 13. doi: 10.1089/ars.2021.0249
Engelbrecht E, Metzler MA, Sandell LL. Retinoid signaling regulates angiogenesis and blood-retinal barrier integrity in neonatal mouse retina. Microcirculation. 2022 Apr; 29(3):e12752. doi: 10.1111/micc.12752. Epub 2022 Mar 5.
Tracy EP, Stielberg V, Rowe G, Benson D, Nunes SS, Hoying JB, Murfee WL, LeBlanc AJ. State of the field: cellular and exosomal therapeutic approaches in vascular regeneration. Am J Physiol Heart Circ Physiol. 2022 Apr 1: 322(4): H647-H680. doi: 10.1152/ajpheart.00674.2021. Epub 2022 Feb 18.
Geller AE, Shrestha R, Woeste MR, Guo H, Hu X, Ding C, Andreeva K, Chariker JH, Zhou M, Tieri D, Watson CT, Mitchell RA, Zhang HG, Li Y, Martin RCG, Rouchka EC, Yan J. The induction of peripheral trained immunity in the pancreas incites anti-tumor activity to control pancreatic cancer progression Nat Commun. 2022 Feb 9;13(1):759. doi: 10.1038/s41467-022-28407-4
Knipe RS, Spinney JJ, Abe EA, Probst CK, Franklin A, Logue A, Giacona F, Drummond M, Griffith J, Brazee PL, Hariri LP, Montesi SB, Black KE, Hla T, Kuo A, Cartier A, Engelbrecht E, Christoffersen C, Shea BS, Tager AM, Medoff BD. Am J Respir Cell Mol Biol. 2022 Jan:66(1):38-52. doi: 10.1165/rcmb.2020-0408OC.
Rowe G, Tracy E, Beare JE, LeBlanc AJ. Cell Therapy rescues aging-induced beta-1 adrenergic receptor and GRK2 dysfunction in the coronary microcirculation. Geroscience. 2022 Feb; 44(1):329-348. doi: 10.1007/s11357-021-00455-6. Epub 2021 Oct 4.
Woolley CR, Brinkman NC, Cash ED, Chandran SK, Mitchell TC. An Unexpected Role for Cell Density Rather than IgM in Cell-Surface Display of the Fc Receptor for IgM in Cell-Surface Display of the Fc Receptor for IgM on Human Lymphocytes. Immunohorizons. 2022 Jan 18: 6(1):47-63. doi: 10.4049/immunohorizons.2100094.
Slomnicki LP, Wei G, Burke DA, Hodges ER, Myers SA, Yarberry CD, Morehouse JR, Whittemore SR, Saraswat Ohri S, Hetman M. Limited Changes in locomotor recovery and unaffected white matter sparing after spinal cord contusion at different times of day. PLoS One. 2021 Nov 23; 16(11):e0249981. doi: 10.137/journal.pone.0249981. eCollection 2021.
Noe JT, Rendon BE, Geller AE, Conroy LR, Morrissey SM, Young LEA, Buntz RC, Kim EJ, Wise-Mitchell A, Barbosa deSouza Rizzo M, Relich ER, Baby BV, Johnson LA,Affronti HC, McMaster KM,Clem BF, Gentry MS, Yan J, Wellen KE, Sun RC, Mitcehll RA. Lactate supports a metabolic-epigenetic link in macrophage polarization. Sci Adv. 2021 Nov 12; 7(46):eabi8602. doi: 10.1126/sciadv.abi8602. Epub 2021 Nov 12
Morrissey SM, Zhang F, Ding C, Montoya-Durango DE, Hu X, Yang C, Wang Z, Yuan F, Fox M, Zhang HG, Guo H, Tieri D, Kong M, Watson CT, Mitchell RA, Zhang X, McMasters KM, Huang J, Yan J. Tumor-derived exosomes drive immunosuppressive macrophages in a pre-metastatic nich through glycolytic dominant metabolic reprogramming. Cell Matab. 2021 Oct 5; 33(10):2040-2058.e10. doi: 10.1016/j.cmet.2021.09.002. Epub 2021 Sep 23.
Tracy EP, Hushes W, Beare JE, Rowe G, Beyer A, LeBlanc AJ. Aging-Induced Impairment of Vascular Function: Mitochondrial Redox Contributions and Physiological/Clinical Implications. Antioxid Redox Signal. 2021 Oct 20; 35(12):974-1015. doi: 10.1089/ars.2021.0031. Epub 2021 Sep 17.
Donninger H, Harrell-Stewart D, Clark GJ. Detection of Endogenous RASSF1A Interacting Proteins. Methods Mol Biol. 2021; 2262:303-310. doi: 10.1007/978-1-0716-1190-6_18.
Morrissey SM, Geller AE, Hu X, Tieri D, Ding C, Klaes CK, Cooke EA, Woeste MR, Martin ZC, Chen O, Bush SE, Zhang HG, Cavallazzi R, Clifford SP, Chen J, Ghare S, Barve SS, Cai L, Kong M, Rouchka EC, McLeish KR, Uriarte SM, Watson CT, Huang J, Yan J. A specific low-density neutrophil population correlates with hypercoagulation and disease severity in hospitalized COVID-19 patients. JCI Insight. 2021 May 10;6(9):e 148435. doi: 10.1172/jci.insight.148435
Wei GZ, Saraswat OS, Khattar NK, Listerman AW, Doyle CH, Andres KR, Karuppagounder SS, Ratan RR, Whittemore SR, Hetman M. Hypoxia-inducible factor prolyl hydroxylase domain (PHD) inhibition after contusive spinal cord injury does not improve locomotor recovery. PLoS One. 2021 Apr 5;16(4):e0249591. doi: 10.1371/journal.pone.0249591. Collection 2021.
Cash E, Sephton S, Woolley C, Elbehi AM, RIA, Ekine-Afolabi B, Kok VC. The role of circadian clock in cancer hallmark acquisition and immune-based cancer therapeutics. J Exp Clin Cancer Res. 2021 Apr 1;40(1):119. doi: 10.1186/s13046-021-01919-5.
Woeste MR, Geller AE, Martin RCG 2nd, Polk HC Jr. Optimizing the Combination of Immunotherapy and Trans-Arterial Locoregional Therapy for Stages B and C Hepatocellular Cancer. Ann Surg Oncol. 2021 Mar;28(3):1499-1510. doi: 10.1245/s10434-020-09414-5. Epub 2021 Jan 3.
Noe JT, Mitchell RA. MIF-Dependent Control of Tumor Immunity. Front Immunol. 2020 Nov 25;11:609948. doi: 10.3389/fimmu.2020.609948. eCollection 2020.
Govindaiah G, Campbell PW, Guido W. Differential Distribution of Ca2+ Channel Subtypes at Retinofugal Synapses. eNeuro. 2020 Nov 5; 7(6):ENEURO.0293-20.2020. doi: 10.1523/ENEURO.0293-20.2020 Print 2020 Nov/Dec.
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University of Louisville MD-PhD Program Mailing Address:
500 Preston Street
HSC Research Tower, Room 315A
Louisville, KY 40202
Parking for Medical School Pre-Orientation Visit: Visitor parking is available at the East Chestnut Street garage, located on the corner of East Chestnut Street and South Preston Street. Medical School Pre-Orientation takes place in February and is located in the Kornhauser Library Auditorium.
Directions: The Health Sciences Center campus is located in the North Central part of Louisville, near the Ohio River.
From the North:
- Take I-65 South to Exit 136 C. Jefferson Street/Downtown/Brook Street.
- Stay in the far left hand lane of the ramp for Brook Street.
- At the end of the ramp, follow Brook Street one block north to East Market Street and turn right.
- Go one block on Market and turn right on South Floyd Street.
- Go four blocks and turn left on East Chestnut Street. The parking garage is on your right at: 414 East Chestnut Street.
From the East or West:
- Take I-64 or I-71 to I-65 South and follow the directions above.
From the South:
- Take I-65 North to Exit 136 A Broadway/Chestnut Street. This ramp exits onto South Brook Street.
- Continue on South Brook Street for two blocks.
- Turn right on East Chestnut Street and continue past South Preston Street. The parking garage is on your right past the pedestrian walkway at: 414 East Chestnut Street.
The University of Louisville (UofL) MD-PhD Summer Undergraduate Research Fellowship (SURF) Program is designed especially for undergraduate students interested in pursuing a combined MD-PhD degree. SURF introduces students to physician-scientist career paths, elucidate the MD-PhD application process and mentor students from socioeconomically disadvantaged backgrounds. Students will spend 10 weeks in the Lab of a UofL faculty mentor, shadow a clinician weekly and have interactions with current MD-PhD Students.
Select Your Area of Research:
- Cancer Biology
- Immunology
- Neuroscience
- Microbiology
- Genetics
- Environmental Disorders
- Health Services and Clinical Investigation
- Bioengineering
- Cardiovascular Biology
- Pharmacology
- Biochemistry
- Physiology
The University of Louisville MD-PhD Summer Undergraduate Research Fellowship Program begins Tuesday, May 26, 2026, and ends Friday, July 31, 2026 - with observance of two University Holidays: Juneteenth (June 19) and Independence Day (July 3).
The 2026 program will accept 10 students. The position requires a full-time effort and commitment. Students accepted into the program will not be able to participate in another activity (e.g., enrollment in classes, prematriculation program, other summer employment, international internship, etc.) during the entire 10 week program.
Program Overview
- 10 week Research Project conducted in the laboratory of a Scientist at the University of Louisville
- Shadowing a clinician a half-day each week
- Weekly Research Seminars
- Interactions with current University of Louisville MD-PhD Students
- Weekly Professional Development Seminars
Program Benefits
- Enhance Research Skills
- Gain insight into MD-PhD application process and physician-scientist careers
- $5,500 stipend allowance
- Housing on the University of Louisville campus
- Travel Allowance
- Networking and Professional Development
Program Requirements
- Complete Research Project
- Attend Weekly Seminars
- Present a 20-minute Presentation and Research Poster at Final Symposium
- Complete Pre- and Post-Evaluation Surveys
Program Eligibility
- You are 18 or older, as of the start of the program
- You are a U.S. Citizen or a Permanent Resident
- You are a Rising Sophomore, Junior or Senior
- You have a 3.0 GPA or higher
Program Information
Rania Gomaa
Program Coordinator Senior
rania.gomaaabdelhay@louisville.edu
Program Application
- Complete the Application: redcap.link/ta2e3ady
- The SURF Application Deadline: Sunday, February 1, 2026
- You will need to write two (2) short papers and provide your educational transcripts, so Apply Early.
The Student Physician-Scientist Accelerated Research and Clinical Training Program (SPARC) is a eight (8) week summer program designed to introduce high school juniors and seniors to the physician-scientist career path by immersing them in hands-on biomedical research, clinical shadowing and professional development. Hosted by the University of Louisville (UofL) MD-PhD Program, this initiative provides early exposure to the intersection of medicine and scientific discovery, equipping students with the knowledge, skills and mentorship to pursue careers as physician-scientists. Students will work on mentored research projects in cutting-edge laboratories at UofL, shadow clinicians and physician-scientists and attend career development workshops focused on leadership, scientific communication and the transition to college and professional training. By the end of the program, students will present their research findings at a scientific symposium, gaining valuable experience in scientific communication.
The Student Physician-Scientist Accelerated Research and Clinical Training Program aims to build a strong pipeline of diverse, highly motivated students who are prepared to enter undergraduate STEM programs, biomedical research careers and physician-scientist training pathways. Through longitudinal mentorship and networking opportunities, SPARC will continue to support students beyond the summer program, fostering a new generation of leaders in medical research and healthcare innovation.
Program Benefits
- Increasing awareness and interest in the physician-scientist career path by providing hands-on exposure to biomedical research and clinical medicine.
- Developing critical research skills by engaging students in laboratory experiences under the mentorship of leading scientists at UofL.
- Enhancing understanding of translational medicine by showing how scientific discoveries impact patient care.
- Providing clinical exposure by shadowing a physician in outpatient clinics, hospitals and specialty areas
- Fostering professional and academic development through workshops on scientific communication, career pathways and college/medical school preparation.
- Preparing students for future academic success by equipping them with the knowledge and skills needed to transition into competitive undergraduate research programs and, ultimately, MD-PhD or MSTP training.
Program Information
- Duration: Summer: 8 weeks
- Eligibility: Applicants who will be high school juniors or seniors in Fall 2026 will be eligible. Applicants must be at least 16 years old by the program’s start date: June 8, 2026. During the program, students must have accommodations in or near Louisville that enable them to participate from 9 a.m. to 5 p.m. via a daily commute. Additionally, they must currently reside in and attend a high school in the U.S. and be U.S. citizens or permanent residents (green card holders) to be eligible to apply.
- Stipend: $2,500
- Final Event: Research Symposium and Awards Poster Presentation
The Student Physician-Scientist Accelerated Research and Clinical Training Program begins Monday, June 8, 2026, and ends Friday, July 31, 2026 - with observance of two University Holidays: Juneteenth (June 19) and Independence Day (July 3).
This 2026 program will accept 8 students. The position requires a full-time effort and commitment. Students accepted into the program will not be able to participate in another activity (e.g., enrollment in classes, prematriculation program, other summer employment, international internship etc.) as a full-time participation is expected (M-F, 9 a.m.-5 p.m.) for the entire eight (8) week program.
Program Contact
Rania Gomaa
Program Coordinator Senior
rania.gomaaabdelhay@louisville.edu
Program Application
- Complete the Application at this website: redcap.link/zo6ulkqj
- The SPARC Application Deadline: Sunday, March 1, 2026
- There are three (3) required documents to provide with your application, so Apply Early.